Chiang Mai University Journal of Natural Sciences

Advisory Boards	Professor Dr. Pongruk Sribanditmongkol, M.D. President Chiang Mai University, Thailand - Associate Professor Dr. Winita Punyodom Vice President Chiang Mai University, Thailand - Assistant Professor Dr. Juthathip Chalermphol Assistant to the President Chiang Mai University, Thailand -
Editor-in-Chief	Associate Professor Dr. Wasu Pathom-aree Faculty of Sciences, Chiang Mai University, Thailand wasu.p@cmu.ac.th
Associate Editors	Professor Dr. Korakot Nganvongpanit Faculty of Veterinary Medicine, Chiang Mai University, Thailand korakot.n@cmu.ac.th Professor Dr. Areewan Klunklin Faculty of Nursing, Panyapiwat Institute of Management, Thailand a.klunklin@gmail.com Distinguished Professor Dr. Anak Iamaroon Faculty of Dentistry, Chiang Mai University, Thailand anak.ia@cmu.ac.th Professor Dr. Pornchai Rachtanapun Faculty of Agro-Industry, Chiang Mai University, Thailand pornchai.r@cmu.ac.th Professor Dr. Supon Ananta Faculty of Science, Chiang Mai University, Thailand supon.a@cmu.ac.th Associate Professor Dr. Veerasak Punyapornwithaya Faculty of Veterinary Medicine, Chiang Mai University, Thailand veerasak.p@cmu.ac.th
Assistant Editors	Associate Professor Dr. Wichai Chattinnawat Faculty of Engineering, Chiang Mai University, Thailand . Associate Professor Dr. Nisit Kittipongpatana Faculty of Pharmacy, Chiang Mai University, Thailand . Associate Professor Dr. Waraporn Boonchieng Faculty of Public Health, Chiang Mai University, Thailand . Associate Professor Dr. Suraphon Lorsomradee, M.D. Faculty of Medicine, Chiang Mai University, Thailand . Assistant Professor Dr. Tonapha Pusadee Faculty of Agriculture, Chiang Mai University, Thailand .
Editorial Boards	Associate Professor Dr. Piyamethd Dilokthornsakul Faculty of Pharmacy, Chiang Mai University, Thailand . Associate Professor Dr. Laongnuan Srisombat Faculty of Science, Chiang Mai University, Thailand . Assistant Professor Dr. Tossapol Moonmanee Faculty of Agriculture, Chiang Mai University, Thailand . Dr. Pachara Sattayawat Faculty of Science, Chiang Mai University, Thailand . Dr. Wipawadee Yooin Faculty of Pharmacy, Chiang Mai University, Thailand . Dr. Pharada Rangseekaew Faculty of Science, Chiang Mai University, Thailand . Dr. Sirasit Srinuanpan Faculty of Science, Chiang Mai University, Thailand . Emeritus Professor Lindsay Falvey Division of Veterinary and Agricultural Sciences, University of Melbourne, Australia . Emeritus Professor Dr. Michael Burgett Collage of Agricultural Sciences, Oregon State University, U.S.A. . Professor Dr. Merilyn Manley-Harris School of Science, University of Waikato, New Zealand . Professor Dr. Keshavan Niranjan Department of Food and Nutritional Sciences, University of Reading, U.K. . Professor Dr. Hichem Frigui School of Engineering, University of Louisville, U.S.A. . Professor Dr. Keung Leung Li Ka Shing Faculty of Medicine, University of Hong Kong, Hong Kong . Professor Manjeet S. Chinnan Department of Food Science and Technology, University of Georgia, U.S.A. . Professor Jeong Sheop Shin Division of Life Sciences, University of Korea, South Korea . Professor Dr. Jonathan V. Sweedler Department of Chemistry, University of Illinois, U.S.A. . Professor Dr. Gary D. Christian Department of Chemistry, University of Washington, U.S.A. . Professor Dr. Kazuyoshi Taya Tokyo University of Agriculture and Technology, Japan . Associate Professor Dr. Nguyen Trong Ngu College of Aquaculture and Fisheries, Can Tho University, Vietnam . Associate Professor Dr. Ian D. McKelvie School of Chemistry, Unversity of Melbourne, Australia . Dr. Janine L. Brown Smithsonian Conservation Biology Institute, U.S.A. . Dr. Lilia de Guzman Agricultural Research Service, United States Department of Agriculture, U.S.A. . Dr. Loganathan Karthik Salem Microbes Private Limited, India -
Managing Editor	Associate Professor Dr. Noppol Leksawasdi Director Office of Research Administration, Chiang Mai University, Thailand .
Assistant Managing Editors	Ms. Priraya Rithaporn Office of Research Administration, Chiang Mai University, Thailand . Mrs. Phatthanan Luesaksiriwattana Office of Research Administration, Chiang Mai University, Thailand . Ms. Thiamchan Panphan Office of Research Administration, Chiang Mai University, Thailand .